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The land snail fauna of several South American countries is very incompletely known despite quite extensive literature. For Bolivia there is only an incomplete and outdated list from 1953, Peru is covered by a name list only (2003), and Ecuador has a catalogue for the mainland Orthalicoidea only (2008). Other countries (Brazil, Chile, Argentina) are covered by recent works from local malacologists, and for some others (Colombia, French Guiana) recently overviews have been published, of which especially the one for French Guiana is also very useful for neighbouring countries like Suriname.
This new paper partly aims to fill that gap for Bolivia, Ecuador and Peru treating the minor families of the Orthalicoidea. Covering the three countries this synopsis summarizes all known species and part of their localities (from literature and some major EU museums). Also one new species is described.
Introduction 5
Methods 6
Systematics 7
Superfamily Orthalicoidea 7
Family Amphibulimidae P. Fischer, 1873 9
Genus Plekocheilus Guilding, 1828 9
Subgenus Plekocheilus (Aeropictus) Weyrauch, 1967 10
Plekocheilus (Aeropictus) tenuissimus Weyrauch, 1967 10
Subgenus Plekocheilus (Eudolichotis) Pilsbry, 1896 11
Plekocheilus (Eudolichotis) hauxwelli (Crosse, 1872) 11
Subgenus Plekocheilus (Eurytus) Albers, 1850 12
Plekocheilus (Eurytus) aristaceus (Crosse, 1869) 14
Plekocheilus (Eurytus) aureonitens (Miller, 1878) 14
Plekocheilus (Eurytus) bruggeni Breure, 1978 15
Plekocheilus (Eurytus) cardinalis (Pfeiffer, 1853) 15
Plekocheilus (Eurytus) doliarius (da Costa, 1898) 16
Plekocheilus (Eurytus) eros (Angas, 1878) 16
Plekocheilus (Eurytus) floccosus (Spix in Wagner, 1827) 17
Plekocheilus (Eurytus) jimenezi (Hidalgo, 1872) 17
Plekocheilus (Eurytus) lynciculus (Deville and Hupé, 1850) 18
Plekocheilus (Eurytus) nocturnus Pilsbry, 1939 19
Plekocheilus (Eurytus) oligostylus Pilsbry, 1939 19
Plekocheilus (Eurytus) onca (d’Orbigny, 1835) 20
Plekocheilus (Eurytus) piperitus piperitus (Sowerby I, 1837) 20
Plekocheilus (Eurytus) piperitus mcgintyi ‘Pilsbry’ H.B. Baker, 1963, stat. n. 21
Plekocheilus (Eurytus) piperitus prodeflexus Pilsbry, 1895, stat. n. 21
Plekocheilus (Eurytus) roseolabrum (E.A. Smith, 1877) 22
Plekocheilus (Eurytus) superstriatus (Sowerby III, 1890) 23
Plekocheilus (Eurytus) taylorianus (Reeve, 1849) 23
Plekocheilus (Eurytus) tricolor (Pfeiffer, 1853) 24
Subgenus Plekocheilus (Plekocheilus) 24
Plekocheilus (Plekocheilus) cecepeus Breure and Araujo, 2015 25
Family Megaspiridae Pilsbry, 1904 25
Genus Paeniscutalus Wurtz, 1947 25
Paeniscutalus crenellus (Philippi, 1867) 26
Genus Thaumastus Martens in Albers, 1860 27
Subgenus Thaumastus (Thaumastiella) Weyrauch, 1956 27
Thaumastus (Thaumastiella) glyptocephalus (Pilsbry, 1897) 28
Thaumastus (Thaumastiella) koepckei Zilch, 1953 28
Thaumastus (Thaumastiella) occidentalis occidentalis Weyrauch, 1960 29
Thaumastus (Thaumastiella) occidentalis debilisculptus Weyrauch, 1960 29
Thaumastus (Thaumastiella) sarcochrous (Pilsbry, 1897) 30
Subgenus Thaumastus (Thaumastus) Albers, 1860 30
Thaumastus (Thaumastus) blanfordianus (Ancey, 1903) 31
Thaumastus (Thaumastus) buckleyi (Higgins, 1872) 31
Thaumastus (Thaumastus) flori (Jousseaume, 1897) 32
Thaumastus (Thaumastus) foveolatus (Reeve, 1849) 32
Thaumastus (Thaumastus) granocinctus (Pilsbry, 1901) 33
Thaumastus (Thaumastus) hartwegi (Pfeiffer in Philippi, 1846) 34
Thaumastus (Thaumastus) inca (d’Orbigny, 1835) 34
Thaumastus (Thaumastus) insolitus (Preston, 1909) 35
Thaumastus (Thaumastus) integer (Pfeiffer, 1855) 35
Thaumastus (Thaumastus) loxostomus (Pfeiffer, 1855) 36
Thaumastus (Thaumastus) magnificus (Grateloup, 1839) 36
Thaumastus (Thaumastus) melanocheilus (Nyst, 1845) 37
Thaumastus (Thaumastus) orcesi Weyrauch, 1967 37
Thaumastus (Thaumastus) orobaenus (d’Orbigny, 1835) 38
Thaumastus (Thaumastus) robertsi robertsi Pilsbry, 1932 38
Thaumastus (Thaumastus) robertsi satipoensis Pilsbry 1944 39
Thaumastus (Thaumastus) sangoae (Tschudi in Troschel, 1852) 39
Thaumastus (Thaumastus) sumaqwayqu sp. n. 40
Thaumastus (Thaumastus) tatutor (Jousseaume, 1887) 41
Thaumastus (Thaumastus) taunaisii (Férussac, 1822) 41
Family Odontostomidae Pilsbry & Vanatta, 1898 42
Genus Cyclodontina Beck, 1837 42
Cyclodontina chuquisacana (Marshall, 1930), comb. n. 42
Cyclodontina lemoinei (Ancey, 1892) 43
Genus Spixia Pilsbry & Vanatta, 1898 43
Spixia minor (d’Orbigny, 1837) 44
Spixia striata (Wagner, 1827) 45
Family Orthalicidae Martens in Albers, 1860 45
Genus Clathrorthalicus Strebel, 1909 45
Clathrorthalicus corydon (Crosse, 1869), comb. n. 46
Clathrorthalicus magnificus (Pfeiffer, 1848) 47
Clathrorthalicus phoebus (Pfeiffer, 1863) 47
Genus Corona Albers, 1850 47
Corona incisa (Hupé, 1857) 49
Corona pfeifferi (Hidalgo, 1869) 50
Corona regalis (Hupé, 1857) 50
Genus Kara Strebel, 1910 51
Kara cadwaladeri (Pilsbry, 1930) 51
Kara indentatus (da Costa, 1901) 52
Kara ortiziana (Haas, 1955) 52
Kara thompsonii (Pfeiffer, 1845) 53
Kara viriata (Morelet, 1863) 53
Kara yanamensis (Morelet, 1863) 54
Genus Orthalicus Beck, 1837 54
Orthalicus bensoni (Reeve, 1849) 55
Orthalicus bifulguratus (Reeve, 1849) 56
Orthalicus mars Pfeiffer, 1861 57
Orthalicus phlogerus (d’Orbigny, 1835) 57
Orthalicus pulchellus (Spix, 1827) 58
Genus Porphyrobaphe Shuttleworth, 1856 58
Subgenus Porphyrobaphe (Oxyorthalicus) Strebel, 1909 59
Porphyrobaphe (Oxyorthalicus) irrorata (Reeve, 1849) 59
Porphyrobaphe (Oxyorthalicus) subirrorata (da Costa, 1898) 60
Subgenus Porphyrobaphe (Porphyrobaphe) Shuttleworth, 1856 60
Porphyrobaphe (Porphyrobaphe) iostoma (Sowerby I, 1824) 60
Porphyrobaphe (Porphyrobaphe) saturnus (Pfeiffer, 1860) 61
Genus Quechua Strebel, 1910 61
Quechua olmosensis olmosensis (Zilch, 1954) 62
Quechua olmosensis maxima (Weyrauch, 1967), stat. n. 62
Quechua salteri (Sowerby III, 1890) 63
Quechua taulisensis (Zilch, 1953) 63
Genus Scholvienia Strebel, 1910 64
Scholvienia alutacea (Reeve, 1850) 64
Scholvienia bambamarcaensis (Breure, 1978) 65
Scholvienia bifasciata (Philippi, 1845) 65
Scholvienia brephoides (d’Orbigny, 1835) 67
Scholvienia claritae (Strebel, 1910) 68
Scholvienia gittenbergerorum (Breure, 1978) 68
Scholvienia huancabambensis Strebel, 1910 69
Scholvienia iserni (Philippi, 1867) 69
Scholvienia jaspidea (Morelet, 1863) 70
Scholvienia jelskii (Lubomirski, 1880) 70
Scholvienia porphyria (Pfeiffer, 1847) 71
Scholvienia weyrauchi (Pilsbry, 1944) 71
Genus Sultana Shuttleworth, 1856 72
Subgenus Sultana (Metorthalicus) Pilsbry, 1899 72
Sultana (Metorthalicus) atramentaria (Pfeiffer, 1855) 73
Sultana (Metorthalicus) augusti (Jousseaume, 1887) 73
Sultana (Metorthalicus) deburghiae (Reeve, 1859) 74
Sultana (Metorthalicus) fraseri (Pfeiffer, 1858) 74
Sultana (Metorthalicus) kellettii (Reeve, 1850) 75
Sultana (Metorthalicus) labeo (Broderip, 1828) 76
Sultana (Metorthalicus) macandrewi Sowerby III, 1889, comb. n. 76
Sultana (Metorthalicus) maranhonensis (Albers, 1854) 77
Sultana (Metorthalicus) shuttleworthi (Albers, 1854) 77
Sultana (Metorthalicus) vicaria (Fulton, 1896) 77
Sultana (Metorthalicus) wrzesniowskii (Lubomirski, 1880) 78
Sultana (Metorthalicus) yatesi yatesi (Pfeiffer, 1855) 78
Sultana (Metorthalicus) yatesi galactostoma (Ancey, 1890) 79
Subgenus Sultana (Sultana) Shuttleworth, 1856 79
Sultana (Sultana) meobambensis (Pfeiffer, 1855), comb. n. 80
Sultana (Sultana) sultana (Dillwyn, 1817) 80
Family Simpulopsidae Schileyko, 1999 81
Genus Simpulopsis Beck, 1837 81
Subgenus Simpulopsis (Eudioptus) Martens in Albers, 1860 82
Simpulopsis (Eudioptus) citrinovitrea (S. Moricand, 1836) 82
Nomen inquirendum 83
‘Pseudoglandina’ agitata Weyrauch, 1967, stat. n. 83
Doubtful and excluded taxa 84
Taxa doubtfully referred to the Ecuadorian malacofauna 84
Taxa doubtfully referred to the Peruvian malacofauna 85
Taxa excluded from the Ecuadorian malacofauna 86
Taxa excluded from the Peruvian malacofauna 86
Discussion 87
Plates 88
Acknowledgements 175
References 175
Appendix 1 186
Appendix 2 188
Appendix 3 190
Appendix 4 192
Appendix 5 195
